It occurs at the surface of warm and warm-temperate seas worldwide; on the Pacific coast of North America it extends from Mexico north to Washington, and it has also been recorded around New Zealand and in the tropical Indian Ocean, the Atlantic and the Mediterranean.
1By-the-Wind Sailor, Velella velella (SIMoN species database)SRC-DSC-EFFF7EDB80
2Velella Velella (or By-the-Wind-Sailors), Point Reyes National SeashoreSRC-DSC-ED65CA6005
3By-the-wind-sailor (Velella velella), MarLIN Marine Life Information NetworkSRC-DSC-C9B974B754
4'By-the-wind sailor' jellies wash ashore in massive numbers after warmer winters (UW News, 18 March 2021)SRC-DSC-37C69DBAA3
5WoRMS REST documented distribution for Velella velella (AphiaID 117832)SRC-DSC-17D4112D61
6Long-term patterns of mass stranding of the colonial cnidarian Velella velella: influence of environmental forcing (Marine Ecology Progress Series 662:69-83, 2021; abstract via Crossref record of DOI 10.3354/meps13644)SRC-DSC-267B3EEA06
7Bluebottle (Australian Museum animal factsheet, updated 15/01/26)SRC-DSC-44A902E837
8Velella velella (Linnaeus, 1758) (World Register of Marine Species taxon page)SRC-DSC-9845085812
9Catalogue of Life (ChecklistBank dataset 3LR) taxon record: Velella velellaSRC-DSC-755FFAD1DC
10GBIF Backbone Taxonomy species record: Velella velella (usageKey 2265793)SRC-DSC-205DA02B0B
Its known predators are few and include nudibranchs of the genus Glaucus, snails of the genus Janthina, the mollusc Fiona pinnata and the fish Mola mola.
SRC-DSC-ED65CA6005 · SRC-DSC-C9B974B754
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These animals float at the surface and are driven ashore by winds, mostly in spring; 20 years of citizen-science data (2000-2019) for the coast from Washington to northern California show mass strandings in 14 years, with especially large events in 2003-2006 and 2014-2019, which researchers link to warmer winters; a university press release states that in the later years carcasses covered more than 1,000 km of continuous coastline within two weeks.